Fossil Homo crania average 723 cubic centimeters before 1.5 million years ago and 1,158 after 600,000 years ago
Source. Du et al., Proceedings of the Royal Society B, 2018, Table S1 (94 crania, CC BY 4.0).
Method and data
Method. Each dot is one cranium: the mean of its published volume estimates, at the midpoint of its date range. The black bars and their numbers are the means of the 21 Homo crania older than 1.5 million years and the 15 younger than 600,000.
| Specimen | Taxon | Group | Endocranial volume (cc, mean of estimates) | Age (million years, midpoint) | Youngest age (million years) | Oldest age (million years) | Region |
|---|---|---|---|---|---|---|---|
| AL 162-28 | Au. afarensis | other | 384 | 3.22 | 3.2 | 3.24 | EA |
| AL 288-1 | Au. afarensis | other | 388 | 3.075 | 2.95 | 3.2 | EA |
| AL 333-105 | Au. afarensis | other | 371 | 3.22 | 3.2 | 3.24 | EA |
| AL 333-45 | Au. afarensis | other | 492 | 3.22 | 3.2 | 3.24 | EA |
| AL 444-2 | Au. afarensis | other | 550 | 3.075 | 2.95 | 3.2 | EA |
| AL 822-1 | Au. afarensis | other | 385 | 3.1 | EA | ||
| Atapuerca 5 | H. heidelbergensis | homo | 1,125 | 0.598 | 0.415 | 0.78 | EU |
| Atapuerca 4 | H. heidelbergensis | homo | 1,390 | 0.598 | 0.415 | 0.78 | EU |
| Bodo 1 | H. heidelbergensis | homo | 1,228 | 0.595 | 0.55 | 0.64 | EA |
| BOU-VP-12/130 | Au. garhi | other | 450 | 2.473 | 2.45 | 2.496 | EA |
| BOU-VP-2/66 | H. erectus | homo | 995 | 0.921 | 0.8 | 1.042 | EA |
| Bukuran | H. erectus | homo | 916 | 1.15 | 0.8 | 1.5 | A |
| Ceprano 1 | H. heidelbergensis | homo | 1,057 | 0.475 | 0.35 | 0.6 | EU |
| D2280 | H. erectus | homo | 783 | 1.765 | 1.76 | 1.77 | EU |
| D2282 | H. erectus | homo | 648 | 1.765 | 1.76 | 1.77 | EU |
| D2700 | H. erectus | homo | 600 | 1.765 | 1.76 | 1.77 | EU |
| D3444 | H. erectus | homo | 638 | 1.765 | 1.76 | 1.77 | EU |
| D4500 | H. erectus | homo | 546 | 1.765 | 1.76 | 1.77 | EU |
| Hexian 1 | H. erectus | homo | 1,025 | 0.412 | A | ||
| KGA-10-525 | P. boisei | other | 545 | 1.42 | 1.41 | 1.43 | EA |
| KNM-ER 13750 | P. boisei | other | 503 | 1.88 | 1.83 | 1.93 | EA |
| KNM-ER 1470 | H. habilis | homo | 756 | 1.959 | 1.868 | 2.05 | EA |
| KNM-ER 1590 | H. habilis | homo | 788 | 1.825 | 1.8 | 1.85 | EA |
| KNM-ER 1805 | H. habilis | homo | 582 | 1.725 | 1.6 | 1.85 | EA |
| KNM-ER 1813 | H. habilis | homo | 496 | 1.725 | 1.55 | 1.9 | EA |
| KNM-ER 23000 | P. aethiopicus | other | 491 | 1.784 | 1.7 | 1.868 | EA |
| KNM-ER 3732 | H. erectus | homo | 738 | 1.907 | 1.868 | 1.945 | EA |
| KNM-ER 3733 | H. erectus | homo | 781 | 1.63 | 1.48 | 1.78 | EA |
| KNM-ER 3883 | H. erectus | homo | 795 | 1.57 | EA | ||
| KNM-ER 406 | P. boisei | other | 500 | 1.63 | 1.56 | 1.7 | EA |
| KNM-ER 407 | P. boisei | other | 474 | 1.85 | EA | ||
| KNM-ER 42700 | H. erectus | homo | 691 | 1.567 | 1.527 | 1.607 | EA |
| KNM-ER 732 | P. boisei | other | 483 | 1.63 | 1.56 | 1.7 | EA |
| KNM-OL 45500 | H. erectus | homo | 715 | 0.882 | 0.79 | 0.974 | EA |
| KNM-WT 15000 | H. erectus | homo | 909 | 1.555 | 1.47 | 1.64 | EA |
| KNM-WT 17000 | P. aethiopicus | other | 410 | 2.526 | EA | ||
| KNM-WT 17400 | P. boisei | other | 430 | 1.77 | 1.67 | 1.87 | EA |
| Lantian | H. erectus | homo | 780 | 1.135 | 1.07 | 1.2 | A |
| MH1 | Au. sediba | other | 420 | 1.977 | SA | ||
| MLD 1 | Au. africanus | other | 510 | 2.715 | 2.58 | 2.85 | SA |
| MLD 37/38 | Au. africanus | other | 445 | 2.58 | SA | ||
| Mojokerto 1 | H. erectus | homo | 800 | 1.62 | 1.43 | 1.81 | A |
| Nanjing 1 | H. erectus | homo | 873 | 0.621 | 0.566 | 0.676 | A |
| Ndutu | H. heidelbergensis | homo | 1,098 | 0.68 | 0.37 | 0.99 | EA |
| Ngandong I | H. erectus | homo | 1,130 | 0.345 | 0.143 | 0.546 | A |
| Ngandong IX | H. erectus | homo | 1,135 | 0.345 | 0.143 | 0.546 | A |
| Ngandong V | H. erectus | homo | 1,281 | 0.345 | 0.143 | 0.546 | A |
| Ngandong VI | H. erectus | homo | 1,097 | 0.345 | 0.143 | 0.546 | A |
| Ngandong VII | H. erectus | homo | 1,047 | 0.345 | 0.143 | 0.546 | A |
| Ngandong X | H. erectus | homo | 1,114 | 0.345 | 0.143 | 0.546 | A |
| Ngandong XI | H. erectus | homo | 1,070 | 0.345 | 0.143 | 0.546 | A |
| Ngandong XII | H. erectus | homo | 1,145 | 0.345 | 0.143 | 0.546 | A |
| OH 12 | H. erectus | homo | 691 | 0.965 | 0.78 | 1.15 | EA |
| OH 13 | H. habilis | homo | 649 | 1.476 | 1.15 | 1.803 | EA |
| OH 16 | H. habilis | homo | 638 | 1.476 | 1.15 | 1.803 | EA |
| OH 24 | H. habilis | homo | 580 | 1.943 | 1.848 | 2.038 | EA |
| OH 5 | P. boisei | other | 509 | 1.844 | 1.839 | 1.848 | EA |
| OH 7 | H. habilis | homo | 685 | 1.844 | 1.839 | 1.848 | EA |
| OH 9 | H. erectus | homo | 1,062 | 1.476 | 1.15 | 1.803 | EA |
| Omo 323-1976-896 | P. boisei | other | 490 | 2.2 | EA | ||
| Omo L.338y-6 | P. aethiopicus | other | 427 | 2.35 | 2.29 | 2.41 | EA |
| Saldanha 1 | H. heidelbergensis | homo | 1,225 | 0.35 | 0.1 | 0.6 | SA |
| Sangiran 10 | H. erectus | homo | 890 | 1.035 | 0.8 | 1.27 | A |
| Sangiran 12 | H. erectus | homo | 963 | 0.935 | 0.8 | 1.07 | A |
| Sangiran 17 | H. erectus | homo | 982 | 0.91 | 0.8 | 1.02 | A |
| Sangiran 2 | H. erectus | homo | 811 | 1 | 0.98 | 1.02 | A |
| Sangiran 3 | H. erectus | homo | 975 | 0.91 | 0.8 | 1.02 | A |
| Sangiran 31 | H. erectus | homo | 700 | 1.7 | 1.6 | 1.8 | A |
| Sangiran 4 | H. erectus | homo | 894 | 1.7 | 1.6 | 1.8 | A |
| Sangiran 38 | H. erectus | homo | 875 | 1.525 | 1.47 | 1.58 | A |
| SK 1585 | P. robustus | other | 494 | 1.98 | 1.6 | 2.36 | SA |
| SK 46 | P. robustus | other | 800 | 1.98 | 1.6 | 2.36 | SA |
| SK 48 | P. robustus | other | 750 | 1.98 | 1.6 | 2.36 | SA |
| SK 54 | P. robustus | other | 500 | 1.98 | 1.6 | 2.36 | SA |
| SK 859 | P. robustus | other | 465 | 1.98 | 1.6 | 2.36 | SA |
| Sts 19/58 | Au. africanus | other | 509 | 2.3 | 2 | 2.6 | SA |
| Sts 5 | Au. africanus | other | 481 | 2.01 | SA | ||
| Sts 60 | Au. africanus | other | 413 | 2.3 | 2 | 2.6 | SA |
| Sts 71 | Au. africanus | other | 442 | 2.3 | 2 | 2.6 | SA |
| Stw 505 | Au. africanus | other | 558 | 2.3 | 2 | 2.6 | SA |
| Taung-1 | Au. africanus | other | 428 | 2.805 | 2.58 | 3.03 | SA |
| Tjg-1993.05 | H. erectus | homo | 858 | 1.15 | 0.8 | 1.5 | A |
| TM 1517 | P. robustus | other | 625 | 1.9 | SA | ||
| Trinil 2 | H. erectus | homo | 901 | 1.535 | 1.47 | 1.6 | A |
| Type 2 | Au. africanus | other | 494 | 2.3 | 2 | 2.6 | SA |
| UA 31 | H. erectus | homo | 995 | 0.992 | EA | ||
| Yunxian | H. erectus | homo | 1,150 | 0.726 | 0.581 | 0.87 | A |
| Zhoukoudian II | H. erectus | homo | 1,013 | 0.627 | 0.504 | 0.75 | A |
| Zhoukoudian III | H. erectus | homo | 915 | 0.627 | 0.504 | 0.75 | A |
| Zhoukoudian V | H. erectus | homo | 1,300 | 0.372 | 0.245 | 0.5 | A |
| Zhoukoudian VI | H. erectus | homo | 850 | 0.602 | 0.423 | 0.78 | A |
| Zhoukoudian X | H. erectus | homo | 1,235 | 0.602 | 0.423 | 0.78 | A |
| Zhoukoudian XI | H. erectus | homo | 1,018 | 0.602 | 0.423 | 0.78 | A |
| Zhoukoudian XII | H. erectus | homo | 1,025 | 0.602 | 0.423 | 0.78 | A |